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Producers Mary Parent, Alex Garca, Eric McLeod and Brian Rogers worked with director Adam Wingard to bring this project to the big screen. The film's script was written by Eric Pearson and Max Borenstein, based on material by Terry Rossio, Michael Dougherty, and Zach Shields. King of the Monsters was written by Borenstein, Dougherty, and Shields as well. In Godzilla vs. King of the Monsters, which takes place five years after the events of King of the Monsters, Godzilla and King Kong face off following Ghidorah's rampage.

Bernie Hayes, an employee of the Apex Cybernetics corporation and the host of the Titan Truth Podcast, a Titan conspiracy theory podcast, exfiltrates information pointing to sinister activities at a Pensacola Apex facility, but just as he is about to finish downloading the files, Godzilla appears and launches an unexpected attack. The remains of the facility include a bizarre piece of technology that resembles a huge robotic eye, which Bernie finds after Godzilla's spree of destruction.

Because Skull Island can no longer hold him (due to Kong's size), Nathan proposes using Kong as their navigator to bring them to the energy source. In the end, Ilene agrees to help halt Godzilla's rampage and locate Kong a new home when Nathan persuades her to do so.

In a modified barge, Nathan, Ilene, Jia, and an Apex crew lead by Walter's daughter Maia board to prevent a mildly drugged Kong from escaping. Kong is locked up to keep him from escaping. Kong gets restless and upset one rainy night, and he begins to tug on the shackles that keep him restrained. Jia calms him down by allowing her and Kong to hold fingers in solidarity as Kong signs the word "home" much to Ilene's surprise. Later, Ilene tells Nathan how she raised Jia as her adopted daughter when violent and unpredictable storms killed most of the Iwi and their parents, but Jia herself was the sole survivor since Kong protected her, resulting in the two of them developing a strong affinity with one another.

When Godzilla is stunned by depth charges, however, their struggle is broken up, allowing Kong to surface for air.

Nathan inquires as to whether Kong would go down the tube if there are others like him if they are unable to return him to Skull Island due to the cold. Ilene responds that he would. Ilene encourages Jia to inquire as to whether Kong's family is present in the area.

Upon finding Bernie, Madison and Josh realize that he is Mark's daughter and he joins them in their investigation. The "eye" Bernie had previously indicated has gone from the Apex headquarters. The group mistakenly sealed and moved Skullcrawler eggs to Hong Kong through an underground conduit.

There, the three disembark and discover Apex's ultimate weapon "Mechagodzilla" a mechanized counterpart of Godzilla being controlled by the late Dr. Ishiro Serizawa's son and Walter's chief technology officer Ren Serizawa. To put Mechagodzilla's strength to the test against the Titans, Walter summons a huge adult Skullcrawler designated "Number 10" into the room as soon as the mecha is ignited. The beast is on the point of attacking Madison, Josh, and Bernie when it is captured and detained by Mechagodzilla, who cleaves the Skullcrawler in two with a tremendous red energy beam, enabling Madison, Josh, and Bernie to take refuge in a tiny hatch. However, the test is cut short as the mecha instantly turns down owing to power supply limits.

The Hollow Earth has an ecosystem that is comparable to Skull Island. Two Warbats attack them, with one of them destroying one of the HEAVs while investigating the area. Kong rescues the group by killing the first Warbat and flinging its corpse onto the second, staggering it for a while before it recovers and constricts itself around him, knocking him out. Nathan's team's HEAV assaulted the Warbat before smothering Kong, enabling Kong to escape and pound the monster to death before eating on its guts.

Following that, Kong and Nathan's squad continue their trek, finally arriving to a massive stone temple that was previously home to Kong's forefathers. They discover remnants of an old fight between Kong's and Godzilla's races as they investigate within, while Kong discovers an ancient axe fashioned from a dead member of Godzilla's race's dorsal plate before sitting on a massive throne. As Nathan's crew discovers the power source necessary to supplement Mechagodzilla's power supply, Kong discovers that the same power source can also be used to replenish the axe. Despite Ilene's protests that Apex cannot just grab the millennium's greatest discovery, Maia and the Apex crew collect the power source and return it to their Hong Kong headquarters using spider-like drones. Maia directs her guards to shoot at Nathan's squad, prompting Kong to aggressively yell at them in defense. Meanwhile, Mechagodzilla starts transmitting a signal to Godzilla, who detects the machine's activation and immediately rushes for Hong Kong.

It's a good thing Godzilla doesn't have the ability to fly. When Maia and her crew attempt to leave their HEAV, Kong pulls it from under their feet and crushes it in his hand, killing Maia and her crew within.

Godzilla and Kong both recuperate and return to their feet, with Godzilla seeking for Kong, who hides from the Titan by perching atop a tower. Kong distracts Godzilla by throwing a construction crane into a building, allowing him to ambush him from above and place the Titan in a chokehold, intending to strangle him, but Godzilla breaks free and unleashes his full fury against Kong, eventually dislocating his left arm, slashing Kong with his claws, and even stomping on his chest twice, despite Kong trying to force Godzilla off of him by punching his wounded thigh. Even though Kong has been conquered and imprisoned by Godzilla, the latter screams defiantly at the latter in defiance.

Godzilla shows mercy and spares his rival's life, as if impressed by Kong's tenacity and prowess, before walking away, leaving Kong alive but badly injured, as he succumbs to commotio cordis.

Walter instructs Ren to send out Mechagodzilla to kill Godzilla. The mecha is activated despite Ren's warnings not to use the new Hollow Earth energy source without testing. Security apprehends Bernie, Madison, and Josh for breaking into Apex HQ and takes them to Walter. While Walter denies inciting Godzilla's war and triggering the whole mess, Madison accuses him of accepting responsibility for providing humanity a chance to fight back against Godzilla and the Titans.

While Josh and Bernie attempt in vain to remotely deactivate the rampaging robot and Madison attempts to summon Mark for assistance, Godzilla and Mechagodzilla battle, at one point both firing their energy beams at each other, which lock together, only for Mechagodzilla to gain the upper hand, knocking Godzilla over and burning him. While Mechagodzilla proceeds to overrun and humiliate Godzilla, Nathan, Ilene, and Jia approach an unconscious Kong, where Jia detects a slowing of his pulse. Ilene explains that they are unable to generate a charge large enough to restart Kong's heart, but Nathan, recalling Maia's earlier statement about the HEAVs producing enough charge to power Las Vegas for a week, flies the remaining HEAV onto Kong's chest and sets it to self-destruct, detonating the craft and releasing a massive electrical charge that instantly revives Kong.

Kong jumps atop Mechagodzilla just as the robot is about to fire its red energy beam into Godzilla's neck, deflecting the beam into the air and sparing Godzilla's life. Godzilla aids Kong in his struggle against Mechagodzilla after realizing Kong saved his life and sense an ally in him after discovering he is not an opponent.

When Godzilla sees Kong wielding his axe, he supercharges it with his atomic breath, enabling Kong to dismember Mechagodzilla limb from limb and pull off its head before hoisting his prize and settling down to rest.

In the aftermath, Nathan, Ilene, and Jia sit together amid the rubble, while Madison reconnects with Mark and introduces him to Josh and Bernie, who is overjoyed to see Mark and invites him onto his podcast to discuss the Roswell Monarch facility. Suddenly, Godzilla approaches Kong, causing the Titan to take his axe and ready to restart the battle. Godzilla, on the other hand, does not attack and rather stares Kong down. Kong lowers his axe, putting an end to the conflict between his species and Godzilla's. Godzilla and Kong see each other as equals after they are no longer competitors, before Godzilla returns to the sea quietly, accompanied by Kong and the assembled people.

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Godzilla vs. Kong has received a mostly favorable reception. As of April 16, 2021, the film has 341 reviews on review aggregation site Rotten Tomatoes, 254 of which are fresh, with a "Tomatometer" rating of 74% certified Fresh. According to the site's reviewers' consensus section, "Delivering squarely on its title, Godzilla vs. Kong swats away character development and human drama to deliver all the spectacle you'd expect from giant monsters slugging it out."

On April 17, 2021, it was announced that Godzilla vs. Kong had eclipsed the overall movie office receipts of Godzilla: King of the Monsters, despite the pandemic's continuing effect on the box office.

This film marks the first time in 59 years that Godzilla and King Kong appeared in a film together since 1962's King Kong vs. Godzilla.

The fact that Chandler appeared in three different Godzilla films, including Godzilla: King of the Monsters and Godzilla vs. Kong, makes him unique in that he is the only individual who has been in a King Kong film, two Godzilla films and one crossover feature between the two.

Neon Genesis Evangelion is mentioned multiple times in the film:

The location where Mechagodzilla is stationed, which is subterranean and topped by a pyramid, is reminiscent of the Central Dogma, which is also underground and topped by a pyramid.